1987 - M.Sc. in Biology. Department of Biology at Kaliningrad State University, USSR
1997 - Ph.D. in Ecology, lichenology. Ekaterinburg Institute of Plant & Animal Ecology, the Ural Division, Russian Academy of Sciences.
2000 - present time - Researcher, Institute of Evolution, University of Haifa.
1991 - 1999 - Researcher, International Center for Environmental Biology at the Tallinn University of Educational Sciences.